Adhikari, S., Timms, W. and Mahmud, M. A. P. (2022): Optimising water holding capacity and hydrophobicity of biochar for soil amendment - A review. Science of The Total Environment, 851(Part 1): 158043. doi: 10.1016/j.scitotenv.2022.158043

Anbazhagan, S., Venugopal, I., Arivazhagan, S., Chinnamuthu, M., Paramasivam, C. R., Nagesh, G., Kannan, S. N., Shamarao, Babu, V. C., Annadurai, M., Muthukkumaran, K., and Rajesh, V. J. (2021): A lunar soil simulant (LSS-ISAC-1) for the lunar exploration programme of the Indian Space Research Organisation. Icarus, 366: 114511. doi: 10.1016/J.ICARUS.2021.114511

Antonangelo, J.A., Culman, S. and Zhang, H. (2024): Comparative analysis and prediction of cation exchange capacity via summation: Influence of biochar type and nutrient ratios. Frontiers in Soil Science, 4: 1371777. doi: 10.3389/fsoil.2024.1371777

Arslan, H., Batiste, S.N. and Sture, S. (2010): Engineering properties of lunar soil simulant JSC-1A. Journal of Aerospace Engineering, 23: 70-83. doi: 10.1061/(ASCE)0893-1321(2010)23:1(70)

Atkin, J., Skabelund, H. A., Pierson, E., Zhen, S., Vandemark, G. and Gentry, T. (2025): Genotype selection and microbial partnerships influence chickpea establishment in lunar regolith simulant. Frontiers in Astronomy and Space Sciences, 12: 1670807. doi: 10.3389/fspas.2025.1670807

Barcenilla, B. B., Kundel, I., Hall, E., Hilty, N., Ulianich, P., Cook, J., Turley, J., Yerram, M., Min, J. H., Castillo-González, C. and Shippen, D. E. (2024): Telomere dynamics and oxidative stress in Arabidopsis grown in lunar regolith simulant. Frontiers in Plant Science, 15: 1351613. doi: 10.3389/fpls.2024.1351613

Barták, M., Hájek, J., Halıcı, M. G., Bednaříková, M., Casanova-Katny, A., Váczi, P., Puhovkin, A., Mishra, K. B. and Giordano, D. (2023): Resistance of primary photosynthesis to photoinhibition in antarctic lichen Xanthoria elegans: Photoprotective mechanisms activated during a short period of high light stress. Plants, 12(12): 2259. doi: 10.3390/plants12122259

Caporale, A., Paradiso, R., Liuzzi, G., Palladino, M., Amitrano, C., Arena, C., Arouna, N., Verrillo, M., Cozzolino, V., de Pascale, S. and Adamo, P. (2023): Green compost amendment improves potato plant performance on Mars regolith simulant as substrate for cultivation in space. Plant Soil, 486: 217-233. doi: 10.1007/s11104-022-05860-0

Coufalík, P., Stavrakakis, H. A. and Argyrou, D. (2024): Chemical analysis of Antarctic regolith and lunar regolith simulant as prospective substrates for experiments with plants. Czech Polar Reports, 14: 241-254. doi: 10.5817/CPR2024-2-15

D'Annunzio, F., Fiscon, G., Conte, F., Paci, P., Bianco, M. D. and Ioio, R. D. (2026): Exploring differential gene correlations in Arabidopsis thaliana for sustainable space farming. Biotechnology Reports (Amsterdam, Netherlands): 49: e00949. doi: 10.1016/j.btre.2026.e00949

Ding, J., Xu, Y., Tan, J., Zhang, H., Xiong, X., Mei, C., Li, M. and Xie, G. (2024): How to make lunar soil suitable for cultivation? - A review. Science of the Total Environment, 948: 174603. doi: 10.1016/j.scitotenv.2024.174603

Duri, L. G., Romano, I., Adamo, P., Rouphael, Y., Pannico, A., Ventorino, V., Pepe, O., De Pascale, S. and Caporale, A. G. (2025): From earth to space: How bacterial consortia and green compost improve lettuce growth on lunar and martian simulants. Biology and Fertility of Soils, 61(7): 1145-1164. doi: 10.1007/s00374-025-01923-3

Genty, B., Briantais, J. M. and Baker, N. R. (1989): The relationship between the quantum yield of photosynthetic electron transport and quenching of chlorophyll fluorescence. Biochimica et Biophysica Acta, 990: 87-92. doi: 10.1016/S0304-4165(89)80016-9

Giordano, D., Barták, M. and Hájek, J. (2025): The combined effect of Cd and high light stress on the photochemical processes in Arabidopsis thaliana. Photosynthetica, 63(2): 182-195. doi: 10.32615/ps.2025.020

Handy, D., Loeffler, A., Knudson, M., Campbell, S., Jaiswal, P., Anderson, J. C. and Goyer, A. (2026): Growth and molecular responses of potato to lunar regolith simulants. bioRxiv. doi: 10.64898/2026.02.23.707481

Haq, S. I. U., Hájek, J. and Barták, M. (2026a): From Antarctic regolith to lunar greenhouses: Mechanistic insights into Brassica rapa photosystem II dynamics for sustainable space agriculture. Life Sciences in Space Research, 48: 111-122. doi: 10.1016/j.lssr.2025.09.008

Haq, S. I. U., Hájek, J., Giordano, D., Mašlaňová, I., Sedláček, I. and Barták, M. (2026b): Psychrotolerant antarctic Mokoshia mucilaginosa and Mokoshia rubra enhance salt stress tolerance in Nicotiana tabacum via photosynthetic stabilization and antioxidant regulation. Plant Cell Reports, 45(3): 61. doi: 10.1007/s00299-026-03732-w

Haq, S. I. U., Hájek, J., Mašlaňová, I., Sedláček, I. and Barták, M. (2025): Antarctic Mokoshia rubra and Mokoshia mucilaginosa enhance growth and cadmium tolerance in Nicotiana species. Plant Physiology and Biochemistry, 229(Part E): 110808. doi: 10.1016/j.plaphy.2025.110808

Holochová, P., Mašlaňová, I., Sedláček, I., Švec, P., Králová, S., Kovařovic, V., Busse, H. J., Staňková, E., Barták, M. and Pantůček, R. (2020): Description of Massilia rubra sp. nov., Massilia aquatica sp. nov., Massilia mucilaginosa sp. nov., Massilia frigida sp. nov., and one Massilia genomospecies isolated from Antarctic streams, lakes and regoliths. Systematic and Applied Microbiology, 43(5): 126112. doi: 10.1016/j.syapm.2020.126112

Krause, H., Weis, W. (1991): Chlorophyll fluorescence and photosynthesis: The basics. Annual Review of Plant Physiology and Plant Molecular Biology, 42: 313-349. doi: 10.1146/annurev.pp.42.060191.001525

Krejčí, Š., Stavrakakis, H.-A., Coufalík, P., Sekerák, J., Hájek, J., Argyrou, D. and Barták, M. (2026): Lunar regolith simulant for cultivation of plants: seed germination and early growth study of plant vitality. International Journal of Astrobiology, 25: e5. doi: 10.1017/S1473550426100275

Lichtenthaler, H. K., Buschmann, C. and Knapp, M. (2005): How to correctly determine the different chlorophyll fluorescence parameters and the chlorophyll fluorescence decrease ratio RFd of leaves with the PAM fluorometer. Photosynthetica, 43(3): 379-393.

Lichtenthaler, H. K., Langsdorf, G., Lenk, S. and Buschmann, C. (2005): Chlorophyll fluorescence imaging of photosynthetic activity with the flash-lamp fluorescence imaging system. Photosynthetica, 43(3): 355-369.

Mei, C., Xie, G. and Wang, X. (2026): Effects of microbial fertilizers on the properties of simulated lunar soil and lettuce growth. Plants, 15(5): 756. doi: 10.3390/plants15050756

Mei, C., Xie, G., Han, Y. and Peng, M. (2025): Biochar derived from plant waste improves seedling growth by influencing the physical structure and nutrient content of simulated lunar soil. Acta Astronautica, 238(Part A): 790-803. doi: 10.1016/j.actaastro.2025.09.033

Müller, P., Li, X. P. and Niyogi, K. K. (2001): Non-photochemical quenching. A response to excess light energy. Plant Physiology, 125(4): 1558-1566. doi: 10.1104/pp.125.4.1558

Ng, C. W. W., Wang, Y. C. (2025): Soil conditioning for enhancing plant growth using biochar and hydrochar under microgravity. NPJ Microgravity, 11(1): 31. doi: 10.1038/s41526-025-00491-y

Ni, J. J., Chen, X. W., Ng, C. W. W. and Guo, H. W. (2018): Effects of biochar on water retention and matric suction of vegetated soil. Geotechnique Letters, 8(2): 124-129. doi: 10.1680/jgele.17.00180

Paul, A. L., Elardo, S. M. and Ferl, R. (2022): Plants grown in Apollo lunar regolith present stress-associated transcriptomes that inform prospects for lunar exploration. Communications Biology, 5(1): 382. doi: 10.1038/s42003-022-03334-8

Qin, L., Yue, Z., Gou, S., Zhang, Y., Wei, G., Shi, K., Zhang, X. and Yang, B. (2025): Structure and formation mechanism of lunar regolith. Space: Science & Technology, 5: 0219. doi: 10.34133/space.0219

Radziemska, M., Gusiatin, M. Z., Cydzik-Kwiatkowska, A., Blazejczyk, A., Holatko, J. and Brtnicky, M. (2023): Does biochar in combination with compost effectively promote phytostabilization of heavy metals in soil under different temperature regimes? Science of the Total Environment, 882: 163634. doi: 10.1016/j.scitotenv.2023.163634

Rombel, A., Krasucka, P. and Oleszczuk, P. (2022): Sustainable biochar-based soil fertilizers and amendments as a new trend in biochar research. Science of The Total Environment, 816: 151588. doi: 10.1016/j.scitotenv.2021.151588

Rosa, A. P., Barão, L., Chambel, L., Cruz, C. and Santana, M. M. (2023): Early identification of plant drought stress responses: Changes in leaf reflectance and plant growth promoting Rhizobacteria selection-the case study of tomato plants. Agronomy, 13(1): 183. doi: 10.3390/agronomy13010183

Shkuratov, Y. G., Bondarenko, N. (2001): Regolith layer thickness mapping of the moon by radar and optical data. Icarus, 149: 329-338. doi: 10.1006/ICAR.2000.6545

Taylor, G. J., Martel, L. M., Lucey, P. G., Gillis-Davis, J. J., Blake, D. F. and Sarrazin, P. C. (2019): Modal analyses of lunar soils by quantitative X-ray diffraction analysis. Geochimica et Cosmochimica Acta, 266: 17-28. doi: 10.1016/J.GCA.2019.07.046

Wamelink, G., Frissel, J., Krijnen, W. and Verwoert, M. (2019): Crop growth and viability of seeds on Mars and Moon soil simulants. Open Agriculture, 4(1): 509-516. doi: 10.1515/opag-2019-0051

Wamelink, G. W., Frissel, J. Y., Krijnen, W. H., Verwoert, M. R. and Goedhart, P. W. (2014): Can plants grow on Mars and the moon: A growth experiment on Mars and moon soil simulants. PLoS One, 9(8): e103138. doi: 10.1371/journal.pone.0103138

Zhang, T., Zhang, T., Chao, C., Yao, Z., Xu, K., Wuxiang, Z., Ding, X., Liu, S., Zhao, Z., Yinghe, A., Yinghe, A., Wang, B., Yu, S., Wang, B. and Chen, H. (2021): The technology of lunar regolith environment construction on Earth. Acta Astronautica, 178: 216-232. doi: 10.1016/j.actaastro.2020.08.039

Zhou, P., Zhao, Z., Wei, G. and Huo, H. (2022): Mineral content estimation of lunar soil of lunar highland and lunar mare based on diagnostic spectral characteristic and partial least squares method. Applied Sciences, 12(3): 1197. doi: 10.3390/app12031197

Web sources / Other sources

[1] Czech Collection of Microorganisms (Brno, Czech Republic): http://www.sci.muni.cz/ccm/